The joint venture (JV) partners have signed an agreement for the FCA to occupy the third stage of the £2.1 billion International Quarter development in Stratford.
The FCA has committed to 425,000 sq ft within the new 515,000 sq ft building.
Transport for London (TfL) previously signed a pre-let agreement for the entirety of a new 265,000 sq ft adjacent building.
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Detailed planning consent has been secured for both buildings and development will being this Summer.
Lend Lease's CEO of International Operations Dan Labbad said: "Securing the FCA as The International Quarter's next major commercial tenant adds to what is quickly becoming London's newest and most progressive business district."
David Joy, Chief Executive at LCR, said: "We are pleased to have reached this pivotal milestone for one of the largest and most significant mixed-use developments in the UK.
"The International Quarter will be a major new employment hub in East London, creating more than 25,000 jobs in total, and we have been working closely with Lend Lease to make this happen."
FCA Acting Chief Operating Officer David Godfrey said: "We look forward to bringing all our London staff together in this exciting new development."
An estimated 6,500 TfL and FCA staff will occupy The International Quarter's offices once the developments are completed in 2017 and 2018.
(LM)
